src=wan,dest=lan would allow any hosts on the internet to connect to any machine on your local network. Perhaps not the best for security and the "option masq 1" in your wan interface allows IP masquerading (known as network address translation) and does all the neccessary config already.
src=lan,dest=lan is the correct one as traffic on the bridge does pass through the firewall on the way through to the other side, and I assume newer versions of the kernel are trying to be flexible by allowing such traffic to also be blocked if you so choose. The only exception to this is two wired clients where the switching is done in hardware.
The reason this is confusing is that OpenWRT's default policy is "block all", with no knowledge of such specific cases like when you are using a bridge. IMO, they should include lan<->lan in the default configs.